Aganane Formation | |
Type: | Geological formation |
Age: | Pliensbachian ~ |
Period: | Pliensbachian |
Prilithology: | Limestone, dolomite |
Otherlithology: | Sandstones, Claystone, Shale, Conglomerate |
Namedfor: | Aganane Village, near Tizouggaghiyn |
Region: | Central High Atlas |
Coordinates: | 31.6°N -6.4°W |
Paleocoordinates: | 25.9°N -4.3°W |
Unitof: | High Atlas |
Underlies: |
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Overlies: |
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Area: | Central High Atlas |
The Aganane Formation is a Pliensbachian (Early Jurassic) geologic formation in the Azilal, Béni-Mellal, Ouarzazate, Tinerhir and Errachidia provinces, central Morocco, being the remnant of a local massive Carbonate platform, and known mostly for its rich tracksites (up to 1350 tracks in 1988) including footprints of thyreophoran, sauropod and theropod dinosaurs.[1] This formation has been dated to the Pliensbachian stage of the Lower Jurassic, thanks to the find of the ammonite Arieticeras cf. algovianum, indicator of Middle Domerian (=Uppermost Pliensbachian) in the upper zone, and lower delimitation by the foraminifers Mayncina termieri and Orbitopsella praecursor (indicators of Carixian=Lower Pliensbachian age).[2] The dinosaur tracksites are all located a few metres below the Pliensbachian-Toarcian limit, being coeval and connected with the lowermost layers of the continental Azilal Formation. The Aganane Formation was also coeval with the Jbel Taguendouft Formation and the Tamadout 1 Formation, all developed along a local "platform-furrow" in the Middle Atlas Mountains, that act as a barrier controlling the western border of the Jurassic Atlas Gulf.[3] The nearshore sections, including both carbonate platforms and close to sea terrestrial facies where located on an isolated internal domain thanks to the control of the barrier, allowing the Aganane Formation to develop on a hot and humid climate, where a local algal marsh had intermittent progradations, intercalated with a layer of terrigenous continental origin.[3] The ichnosites were developed in tidal flats and coastal deposits suitable to sea flooding.[4]
The Aganane Formation is a member of the Pliensbachian facies section of the Central Atlas, which are distributed from west to east: The Aït Chitachen Formation & Aït-Bazzi Formation at Demnate (continental-fluvial, coastal lagoon), the Aganane itself at Azilal.[4] At Tazoult, part of the Azilal profile contacts the bottom with the karst Talmest-Tazoult Formation, then a section where the Aganane itself indicates an eastward expansion of the carbonate facies, finally, the most recent Pliensbachian strata belong to the Amezraï Formation (Intertidal to coastal marine), this last change being the result of a transgression where we see a westward advance of the Imilchil pelagic facies.[4]
At Tazoult, the presence of a Diapir (the Tazoult salt wall) is remarkable. In the Pliensbachian, this area begins with the deposition of the Talmest-Tazoult Formation, and then is invaded from the west by the shallow marine carbonate platform of the Aganane Formation.[5] At this time, a decrease in the growth rate of the diapir is detected compared to the Sinemurian. In fact, with the eastward extension of the Aganane Formation, the Tazoult salt wall registers a major change towards shallower facies, confirming a higher diapir relief and a decrease in water depth towards this area during the Pliensbachian, not excluding a complete stop of diapir activity in this interval.[5]
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During the Pliensbachian epoch, the region was situated at a nearly tropical paleolatitude and marked the western boundary of the ancient "Atlas Gulf," which faced the Tethys Sea to the East. To the north of the Demnate fault, there existed an extensive subsiding tidal flat where carbonate deposits containing gypsum remnants and stromatolitic laminites formed, interspersed with desiccation polygons.[6] Along the fault's edge, active during this period, lignite layers were intercalated, likely resulting from the degradation of forests located to the south, indicated by root traces in sandstones at the base of the Aït-Bazzi Formation in Ait Tioutline. The area was inhabited by both herbivorous and carnivorous dinosaurs that roamed the expansive coastal marshland. Towards the east, excluding the Demnat region, gypsum precipitated in sebkhas along the edge of a low continent, indicative of a possibly arid climate. Additionally, more massive facies with large lamellibranchs separated the tidal flat from the open sea with Ammonites, whose western extension bordered the eastern edge of Azilal.
The Aganane Formation is characterized as representing a carbonate platform, the most important Lower Jurassic platform in the central High Atlas, bounded to the west at Demnat by the Aït Chitachen Formation, which represents environments ranging from fluvial to coastal lagoons, with palynological residues Corollina quezelii & Corollina yvesi (Cheirolepidiaceae) as dominant, indicating an arid environment, accompanied by Cupressacites oxycedroides & Diadocupressacites moghrebiensis, both palynological residues of Cupressaceae.[7]
The Carbonates of this unit were subjected to an examination of the diagenetic characteristics of the carbonates, allowing the construction of a model of diagenetic environments, where changes in the surface environment were consistently reflected in a sequence of rocks by diagenetic characteristics, maybe a tentative correlation of major events, such as Hurricanes.[8] The carbonate rocks that form the massive "Aganane Platform" are clearly influenced by tidal and shallow marine flows, ascribed to three environments: supratidal, intertidal and subtidal.[9] The supratidal sector is diverse, including continental deposits with abundant quartz-rich cricundant deposits and fluvial channels, as well as others composed of considerable thicknesses of stratified gypsum and chicken-wire cargneules, lime and dolomite shales and marls, with drying cracks, caliche crusts and vadose pisoliths. These sediments suggest a continental zone bordering river courses, connected to coastal sectors where shales and siltstones derived from Sabkhas in which interstitial evaporites were formed.[10] [11] Intertidal deposits are composed of algal laminates, bioturbated pelletiferous shales, uniformitarian shales and wackestones with disruptive channels and storm sequences. Algal laminated Boundstones developed in both supratidal and intertidal zones, following present-day patterns such as Shark Bay in Australia or the Persian Gulf. Whereas the pelletoid lime packstone/wackestone facies most likely represented tidal flat areas, or analogs to modern coastal mangrove growth.[10] [11] Subtidal deposits include both those of an intertidal nature, indicating shallow water or coastal lagoons, with extensive accumulations of skeletal lime packstones, oolitic tidal deltas and offshore bars, oncoliths and coral reefs, and occasional Opisoma spp. While further east the more open subtidal conditions are represented by flint-bearing calcareous shales and sparse faunas where ammonites begin to appear.[10] [11]
The Carbonate Facies of the High Atlas "Middle Lias" (Pliensbachian) generally have a low bathymetry. They show a notable evolution from east to west from a lagoon-marine facies to brackish facies. At the level of the Azilal Atlas, the Middle Lias occurs on both sides of the Demnat Fault: to the southwest of the fault, the Aït Bazzi Formation with dolomitic and red marly facies (lagoonal and lagoon-evaporitic facies), while to the northeast develop the thicker facies of the Aganane Formation.[12] In this area, formations of this stage begin either with conglomeratic facies associated with red marls or with scree and conglomerates associated with local erosion, recovering the same lower Pliensbachian emersion phase observed in the Beni-Mellal Atlas.[12] Toward the south, the "basin" facies ends at the bottom of the gulf near the Jbel Oukarde accident, at the western end of the Tilougguit syncline. The southern limit of this basin, hidden by the later sediments, corresponds to the present-day course of the Azilal-Anergui submeridian fault. Toward the south, on the Amezraï and Aït Bouguemmez basins, a platform zone was established in the heart of the Central High Atlas, consisting of the Jbel Choucht, Aganane, Assemsouk, and Amezraï formations.[12] The Jbel Choucht Formation is the type locality here, characterized by a great development of coralligenous facies. The Jbel Choucht Formation is also present towards the north-central High Atlas, but without any reef character, being rather rich in megalodont bivalves, while the Aganane Formation presents more or less the same characteristics as those described in the Beni-Mellal Platform. To the east, at the level of the Jbel Aroudane, the thickening and sedimentary polarity of the Jbel Choucht Formation occurs this time southward, with a W-E subsident basin centered on the Jbel Azourki and the Jbel Aroudane.[12]
In the Aganane Formation towards terminal Carixian (Lias biozone C2 in the High Atlas of Morocco) several sedimentary environments are distinguished in the carbonate inner platform. These environments are characterized by rhythmic sedimentation composed of metric sedimentary sequences analogous to present-day "marnières".[13] Lithologically, there are two juxtaposed sedimentary units that represent two types of complex environments: a first assemblage that includes the marlodolomitic unit of the Aganane Formation and the Aït Bazzi Formation; these units are also equivalent to lagoonal and lagoon-evaporitic facies.[13] This sedimentary assemblage corresponds to a carbonate supratidal coastal plain environment regularly overlain by red marl deposits of continental origin, which can be compared to the modern Sabkhas in the Persian Gulf, but in a less arid climatic context, for the frequent presence of dinosaur tracks suggests extensive vegetation cover in the hinterland, and thus high annual precipitation, suggesting overall tropical conditions reminiscent of the Andros Island model in the Bahamas.[13] A second set (a light gray limestone-dolomite subunit) more distal to the inner platform, in which the environment varies, repeatedly, from subtidal to supratidal conditions, from marine to coastal plain deposits, sometimes showing red terrigenous intercalations, revealing the presence of local cycles evolving from temporary lagoon to "sabkha", while in the outermost part of the platform the existence of a permanent lagoon can be assumed.[14] The facies of this lagoon are muddy and highly bioturbated, with numerous marine organisms colonizing the soft substrate of the lagoon: Lamellibranchs, Gastropods, Brachiopods, with calcareous algae (Palaeodasycladus, Solenopores, etc.) oncoliths and Foraminifers. These levels with large Lamellibranchs (especially "Lithiotidae") are rather rare in the Carixian.[13]
Locations such as Ait Athmane recover the typical Sinemurian-Pliensbachian mediterranean lithiotid bivalve reefs, composed by aggrupation of aberrant bivalves.[15] These "Reefs" had a strong zonation, starting with the bivalves Gervilleioperna and Mytiloperna, restricted to intertidal and shallow-subtidal facies. Lithioperna is limited to lagoonal subtidal facies and even in some low-oxygen environments. Finally Lithiotis and Cochlearites are found in subtidal facies, constructing buildups.[16] Locally, these reefs were developed as shallow subtidal, cross-bedded floatstones, later evolving to layers with evidence of subaerial exposure, including lagoonal marls, and bioturbated red mudstones with root traces and calcrete.[15] These layers are abundant on the aberrant bivalves Lithioperna and Cochlearites, as well common corals, gastropods, the bivalve Opisoma and oncoids, all living in a sheltered lagoon in the interior of the local carbonate platform, similar to the Rotzo Formation of the Trento Platform.[15]
At Ait Bou Guemmez, the upper Aganane Formation records the development of a lagoonal environment south of the Jbel Tizal-Jbel Azourki accident, which evolves to a more or less open subtidal platform environment north of this accident. The local transgressive procession is marked by the dominance of subtidal facies with a microfauna of biozone C1, whereas the high marine level procession is represented by a succession of intertidal to supratidal dolomitic limestones, its top is surmounted by an important subaerial discontinuity materialized by red clay with paleosols and abundant plant roots (e.g. at Tizi n'Terghist). In other localities are mainly biodetritic limestones, showing emersion structures at their top (dolomitization, mud cracks, roots and plant remains, footprints of Dinosaurs).[17]
The Aganane Formation is located largely within the "Grand e Accident du Nor d'Altasien" or North Atlas Fault, the largest and most important of the Central High Atlas faults.[11] During the late Pliensbachian, 200 m of carbonate sediments accumulated south of the fault on an overturned block, while to the north more than 700 m of similar material accumulated.[11] This fault line probably marked the northern boundary of a Paleozoic basement peninsula that advanced eastward from the Tichka Massif into the Atlas Trench.[11]
The paleogeographic evolution of our region can be described in three main stages:
Genus | Species | Stratigraphic position | Material | Habitat | Notes | Images |
---|---|---|---|---|---|---|
Everticyclammina[18] |
|
| Calcareous Skeletons | Marine or Lagoonal | A foraminifer of the Everticyclamminidae family. It represents a species similar to E. virguliana, known from the Middle Jurassic of Morocco. | |
Haurania |
|
| Calcareous Skeletons | Marine or Lagoonal | A foraminifer of the Hauraniinae family. | |
Lituosepta |
|
| Calcareous Skeletons | Marine or Lagoonal | A foraminifer of the Mesoendothyridae family. | |
Mayncina |
|
| Calcareous Skeletons | Marine or Lagoonal | A foraminifer of the Nezzazatoidea family. | |
Mesoendothyra |
|
| Calcareous Skeletons | Marine or Lagoonal | A foraminifer of the Mesoendothyridae family. | |
Orbitopsella |
|
| Calcareous Skeletons | Marine or Lagoonal | A foraminifer of the Mesoendothyridae family. | |
Pseudocyclammina |
|
| Calcareous Skeletons | Marine or Lagoonal | A foraminifer of the Pfenderinidae family. | |
Pseudopfenderina |
|
| Calcareous Skeletons | Marine or Lagoonal | A foraminifer of the Pfenderinidae family. | |
Siphovalvulina |
|
| Calcareous Skeletons | Marine or Lagoonal | A foraminifer of the Pfenderinidae family. | |
Genus | Species | Stratigraphic position | Material | Habitat | Notes | Images |
---|---|---|---|---|---|---|
Boueina |
|
| Calcareous Skeletons | Marine or Lagoonal | An alga of the Halimedaceae family. | |
Cayeuxia |
|
| Calcareous Skeletons | Marine or Lagoonal | An alga of the Halimedaceae family. | |
Paleodasycladus |
|
| Calcareous Skeletons | Marine or Lagoonal | An alga of the Dasycladaceae family. | |
Pseudolithocodium |
|
| Calcareous Skeletons | Marine or Lagoonal | An alga of the Thaumatoporellales family. | |
Sestrosphera |
|
| Calcareous Skeletons | Marine or Lagoonal | An alga of the Solenoporaceae family. This genus is the most common algal type found in the Biozone A (Lituosepta recoarensis). | |
Thaumatoporella |
|
| Calcareous Skeletons | Marine or Lagoonal | An alga of the Thaumatoporellales family. | |
Genus | Species | Location | Material | Type | Made by | Notes | Images |
---|---|---|---|---|---|---|---|
Arenicolites |
|
| Borrowing Traces | Domichnia | Marine, brackish, or freshwater Unbranched U-shaped burrows, subvertical in orientation, with or without lining and passive infill. Are common in modern coastal environments. | ||
Chomatichnus |
| Tubular Fodinichnia | Fodinichnia |
| Interpreted as the feeding burrow of a sediment-feeding animal. | ||
Chondrites |
| Tubular Fodinichnia | Fodinichnia |
| Burrow-like ichnofossils. Interpreted as the feeding burrow of a sediment-ingesting animal. A more recent study has found that Scoloplos armiger and Heteromastus filiformis, occurring in the German Wadden Sea in the lower parts of tidal flats, make burrows that are homonymous with numerous trace fossils of the ichnogenus.[19] | ||
Rhizocorallium |
| Tubular Fodinichnia | Domichnia and/or fodinichnia. |
| Dwelling and feeding burrow of a suspension-feeder or deposit-feeder, associated usually with shallow waters | ||
Skolithos |
| Cylindrical to subcylindrical burrows | Domichnia |
| Burrow-like ichnofossils made by organisms advancing along the bottom surface. Very narrow, vertical or subvertical, slightly winding unlined shafts filled with mud. Interpreted as dwelling structures of vermiform animals; specifically, the domichnion of a suspension-feeding worm or phoronidans | ||
Thalassinoides |
| Tubular Fodinichnia | Fodinichnia |
| Burrow-like ichnofossils. Large burrow-systems consisting of smooth-walled, essentially cylindrical components. Common sedimentary features are Thalassinoides trace fossils in the fissile marlstone to claystone intervals | ||
Zoophycos |
| Dwelling traces | Domichnia & Fodinichnia |
| Burrow-like ichnofossils. It has been related to Echiurans, but also from moving and feeding polychaete worms. | ||
Genus | Species | Stratigraphic Position | Material | Notes | Images |
---|---|---|---|---|---|
Ampakabastraea |
|
| Calcified skeleton pieces | A Coral of the family Stylinidae. | |
Archaeosmilia |
|
| Calcified skeleton pieces | A Coral of the family Zardinophyllidae. Late Pliensbachian corals of the Aganane Formation are mainly limited to Retiophyllia, Thamnasteria and Archaeosmilia. | |
Archaeosmiliopsis |
|
| Calcified skeleton pieces | A Coral of the family Archaeosmiliidae. | |
Oppelismilia[20] |
|
| Calcified skeleton pieces | A Coral of the family Oppelismiliidae. | |
Phacelostylophyllum |
|
| Calcified skeleton pieces | A Coral of the family Stylophyllidae. | |
Phacelophyllia |
|
| Calcified skeleton pieces | A Coral of the family Dermosmiliidae. | |
Periseris |
|
| Calcified skeleton pieces | A Coral of the family Latomeandridae. | |
Retiophyllia |
|
| Calcified skeleton pieces | A coral of the family Reimaniphylliidae. The phaceloid genus at this site is similar to the common Triassic genus Retiophyllia. | |
Thamnasteria |
|
| Calcified skeleton pieces | A coral of the family Thamnasteriidae. | |
Genus | Species | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|
Calcirhynchia[21] |
|
| Isolated Shells | A Brachiopodan of the family Cirpinae. Relatively abundant on seashore deposits. It was originally identified as part of the genus Rhynchonella | |
Hesperithyris |
|
| Isolated Shells | A Brachiopodan of the family Zeilleriidae | |
Spiriferina |
|
| Isolated Shells | A Brachiopodan of the family Spiriferinidae | |
Tauromenia |
|
| Isolated Shells | A Brachiopodan of the family Zeilleriidae | |
Genus | Species | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|
Arcomytilus |
|
| Isolated Shells | A saltwater bivalve of the family Mytilidae. | |
Cochlearites[22] [23] |
|
| Isolated Shells | A saltwater bivalve of the family Pterioida. A large bivalve, with a subequal shell, that can reach 60–70 cm in height. It is one of the three principal bivalves found on the Lithiotis facies, whose accumulations generally cover megalodontid coquinas. | |
Eomiodon |
|
| Isolated Shells | A saltwater/brackish bivalve of the family Neomiodontidae. This genus is considered an opportunistic suspension feeder of shallow infauna, and the marker genus for brackish environments.[24] | |
Fimbria |
|
| Isolated Shells | A saltwater bivalve of the family Lucinidae. | |
Gervilleia |
|
| Isolated Shells | A saltwater bivalve of the family Bakevelliidae. | |
Gervillioperna |
|
| Isolated Shells | A saltwater bivalve of the family Malleidae. Abundant along the rootlets, indicating a very shallow and restricted lagoon or swamp environment | |
Liogryphaea |
|
| Isolated Shells | A saltwater bivalve of the family Gryphaeidae. This genus develops a noted material oyster biostrome at Aït Athmane, where a discontinuous, patchy layer is formed, developed under submarine lithification and a relative enrichment in terrigenous matter. | |
Lithioperna |
|
| Isolated Shells | A saltwater bivalve of the family Pterioida. This genus was founded to be a bivalve with a juvenile byssate stage that developed different lifestyles in adulthood depending on the density of individuals and the firmness of the bottom | |
Lucina |
|
| Isolated Shells | A saltwater/brackish bivalve of the family Lucinidae. Linked with intertidal settings | |
Mytiloperna |
|
| Isolated Shells | ||
Opisoma[25] |
|
| Isolated Shells | A saltwater/brackish bivalve of the family Astartidae. Is considered a genus that evolved from shallow burrowing ancestors, secondarily becoming an edge-lying semi-fauna adapted to photosymbiosis. In the Aganane Formation, this genus is both associated with Corallinaceous facies and tidal flats, even recovered on intertidal channels similar to the ones recovered on modern mangroves | |
Pachyrisma[26] |
|
| Isolated Shells | A saltwater bivalve of the family Pachyrismatidae | |
Pecten |
|
| Isolated Shells | ||
Pholadomya |
|
| Isolated Shells | A saltwater bivalve of the family Pholadomyidae | |
Protodiceras |
|
| Isolated Shells | A saltwater bivalve of the family Pachyrismatidae | |
Genus | Species | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|
Nerinea |
|
| Isolated Shells | A saltwater gastropod of the family Nerineidae. Local specimens appear to have algal material on the shells, indicating a restricted lagoon environment. | |
Pseudonerinea |
|
| Isolated Shells | A saltwater gastropod of the family Nerineidae | |
Scurria |
|
| Isolated Shells | A saltwater gastropod of the family Nerineidae. "Scurria" was found in the Assemsouk structure on a "Cochlearites" valve and a shallow ovoid excavation, similar to the resting trace of a limpet, was found inside a transported "Lithiotis". | |
Genus | Species | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|
Arieticeras |
|
| Isolated Shells | An ammonite of the family Hildoceratidae. Arieticeras cf. algovianum is indicative of the Middle Domerian (Upper Pliensbachian) in the upper zone | |
Fuciniceras |
|
| Isolated Shells | An ammonite of the family Hildoceratidae | |
Seguenziceras |
|
| Isolated Shells | An ammonite of the family Polymorphitidae | |
Tropidoceras |
|
| Isolated Shells | An ammonite of the family Polymorphitidae | |
Uptonia |
|
| Isolated Shells | An ammonite of the family Polymorphitidae | |
Genus | Species | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|
Anchisauripus |
|
| Footprints | ||
Argoides[27] |
|
| Footprints | Theropod tracks, member of the ichnofamily Anchisauripodidae, incertae sedis inside Neotheropoda. Mistaken originally as coelurosaur tracks, this specimens have pes that resemble those of Noasaurids and other Abelisauroids.[28] Includes a pathologic trackway with evidence of limping, as well three morphotypes: Didactyl, tridactyl and tetradactyl theropod trackways. | |
Carmelopodus |
|
| Footprints | Theropod tracks, member of the ichnofamily Eubrontidae, incertae sedis inside Theropoda. Ceratosaur tracks. Includes the largest theropod track from the Early Jurassic of Morocco. | |
Eubrontes[29] |
|
| Footprints | Theropod tracks, type member of the ichnofamily Eubrontidae, incertae sedis inside Theropoda. Eubrontes is related to the Genus Dilophosaurus, representing a basal Neotheropods. | |
Grallator |
|
| Footprints | Theropod tracks, typical member of the ichnofamily Grallatoridae, incertae sedis within Theropoda. Attributed to dinosaurs of type Coelophysidae and Dilophosaurus | |
Megalosauripus |
|
| Footprints | Traces of theropods, members of the ichnofiber family Eubrontidae, incertae sedis within Theropoda. Traces of tetanurans, some with a resemblance to Allosauroids. | |
Plesiornis? |
|
| Footprints | Traces of theropods, members of the ichnofamily Anchisauripodidae, incertae sedis within Neotheropoda. Theropods with bird-like legs or similar to that of Coelurosaurs | |
Theropodipedia[30] [31] | Indeterminate |
| Footprints | Theropod Tracks of uncertain affinity | |
Genus | Species | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|
Breviparopus |
|
| Footprints | Traces of sauropods, typical member of the ichnofamily Parabrontopodidae, incertae sedis within Sauropodomorpha. Includes traces with pes similar to Diplodocoidea.[32] | |
Eosauropus |
|
| Footprints | Traces of sauropods, typical member of the ichnofamily Parabrontopodidae, incertae sedis within Sauropodomorpha. Includes traces with pes similar to those of Blikanasaurus. | |
Lavinipes?[33] |
|
| Footprints | Traces of sauropods, typical member of the ichnofamily Parabrontopodidae, incertae sedis within Sauropodomorpha. | |
Otozoum[34] |
|
| Footprints | Traces of sauropodomorphs, a type member of the ichnofamily Otozoidae, incertae sedis within Sauropodomorpha. Includes a gigantic 84 or 75 cm track that represents the largest Otozoum ever described in the literature. | |
Parabrontopodus[35] |
|
| Footprints | Traces of sauropods, typical member of the ichnofamily Parabrontopodidae, incertae sedis within Sauropodomorpha. Includes tracks with pes similar to those of Vulcanodon (perhaps left by Tazoudasaurus?) and other morphotype more similar to Rhoetosaurus. | |
Pseudotetrasauropus |
|
| Footprints | Traces of sauropodomorphs, a type member of the ichnofamily Otozoidae, incertae sedis within Sauropodomorpha. | |
Sauropodomorphidia[36] | Indeterminate |
| Footprints | Sauropodomorph tracks of uncertain affinity | |
Sauropodina | Indeterminate |
| Footprints | Sauropod tracks of uncertain affinity | |